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LC
Tufted Duck Aythya fuligula

Justification
This species has an extremely large range, and hence does not approach the thresholds for Vulnerable under the range size criterion (Extent of Occurrence <20,000 km2 combined with a declining or fluctuating range size, habitat extent/quality, or population size and a small number of locations or severe fragmentation). The population trend appears to be stable, and hence the species does not approach the thresholds for Vulnerable under the population trend criterion (>30% decline over ten years or three generations). The population size is extremely large, and hence does not approach the thresholds for Vulnerable under the population size criterion (<10,000 mature individuals with a continuing decline estimated to be >10% in ten years or three generations, or with a specified population structure). For these reasons the species is evaluated as Least Concern.

Taxonomic source(s)
AERC TAC. 2003. AERC TAC Checklist of bird taxa occurring in Western Palearctic region, 15th Draft. Available at: #http://www.aerc.eu/DOCS/Bird_taxa_of _the_WP15.xls#.
Cramp, S.; Perrins, C. M. 1977-1994. Handbook of the birds of Europe, the Middle East and Africa. The birds of the western Palearctic. Oxford University Press, Oxford.
del Hoyo, J.; Collar, N. J.; Christie, D. A.; Elliott, A.; Fishpool, L. D. C. 2014. HBW and BirdLife International Illustrated Checklist of the Birds of the World. Barcelona, Spain and Cambridge UK: Lynx Edicions and BirdLife International.

Population justification
The global population is estimated to number c.2,600,000-2,900,000 individuals (Wetlands International 2006), while national population sizes have been estimated at c.10,000-100,000 breeding pairs and c.1,000-10,000 wintering individuals in China and c.1,000-10,000 individuals on migration and c.1,000-10,000 wintering individuals in Taiwan (Brazil 2009).

Trend justification
The overall population trend is stable, although some populations have unknown trends (Wetlands International 2006).

Ecology
Behaviour In central and north-west Europe this species is mainly sedentary (del Hoyo et al. 1992) although other populations are chiefly migratory (Madge and Burn 1988). Migratory populations arrive on their breeding grounds from late-April (Kear 2005a) where they breed in single pairs or loose groups (del Hoyo et al. 1992) with hundreds often nesting at the same site (although not colonially) (Kear 2005a). Once incubation has commenced (Madge and Burn 1988) the males gather in flocks (Madge and Burn 1988, Kear 2005a) to moult (moult migrations occurring in some areas) between late-June and early-September when they become flightless for 3-4 weeks (Scott and Rose 1996) (females moult their flight feather 1-2 months later) (Scott and Rose 1996). The autumn migration begins in September (Kear 2005a), with a segregation of the sexes occurring in autumn and winter due to the difference in the timing of the moult (Scott and Rose 1996). The return spring migration begins in late-February (Kear 2005a). During the winter the species is highly gregarious and may occur in flocks of several thousand individuals (Kear 2005a). Habitat Breeding The species breeds in lowland regions and shows a preference for eutrophic waters 3-5 m deep (avoiding lakes deeper than 15 m) (Kear 2005a) with open water, islands for breeding (del Hoyo et al. 1992) and abundant marginal and emergent vegetation (Kear 2005a). It is common on large, freshwater lakes, ponds, reservoirs (del Hoyo et al. 1992, Kear 2005a), gravel-pits (Kear 2005a) and quiet stretches (Kear 2005a) of wide slow-flowing rivers during this season (del Hoyo et al. 1992, Kear 2005a). Non-breeding During the winter the species frequents large freshwater lakes (Kear 2005a), reservoirs (Snow and Perrins 1998) and sheltered coastal locations (Kear 2005a) such as brackish lagoons (del Hoyo et al. 1992), brackish inland seas (Kear 2005a) (e.g. Caspian Sea) (Scott and Rose 1996), tidal bays (del Hoyo et al. 1992) and estuaries (Madge and Burn 1988, Snow and Perrins 1998) although it avoids strong wave action and very exposed maritime conditions unless all inland freshwaters become frozen (Snow and Perrins 1998). Diet The species is omnivorous (Kear 2005a) a major part of its diet consisting of molluscs (especially Mytillus and Cardium spp., gastropods (del Hoyo et al. 1992) and zebra mussels Dreissena polymorph (Kear 2005a)), crustaceans and aquatic insects (del Hoyo et al. 1992), as well as grain (Kear 2005a) and the seeds and vegetative parts of aquatic plants (del Hoyo et al. 1992). Breeding site The nest is constructed of vegetation and is placed in water (Kear 2005a) on floating mats or islets (Flint et al. 1984), or on the ground on islands (del Hoyo et al. 1992) in rush or grass tussocks (Snow and Perrins 1998), under bushes (Kear 2005a) or in the open amidst marsh-nesting gull or tern colonies (for protection against predators) (Madge and Burn 1988, Kear 2005a). Nests are usually placed within 20 m of water (Kear 2005a) although on islands they may be placed up to 150 m away (Snow and Perrins 1998). Although the species is not colonial hundreds may nest on the same site with neighbouring nests spaced 7-11 m apart (within gull or tern colonies nests are spaced 2-3 m apart) (Kear 2005a). Management information In the UK (Salford docks, Manchester) the species prefers to feed in waters heavily polluted with sewage that are devoid of aquatic vegetation but hold high densities of oligocheates and other pollution-tolerant organisms (Marsden and Bellamy 2000). The species may therefore suffer from plans to improve water quality in the docks (e.g. modernising sewage treatment systems and oxygenating water) (Marsden and Bellamy 2000). Experimental removal (extermination) of the nest predator American mink Neovison vison in the outer archipelago of south-west Finland also resulted in an increase in the breeding density of this species (Nordstrom et al. 2002). The species competes for food (e.g. chironomid larvae and molluscs) with fish species such as roach Rutilus rutilus (Winfield and Winfield 1994). A study in the Czech Republic found that fish ponds with a fish stock density of less than 400 kg ha1, water transparency of more than 50 cm, mixed fish stocks (e.g. tench and pike or perch) rather than monospecific stocks (e.g. of carp), and systems that include ponds with fish fry (to provide areas with low fish competition and high invertebrate availability) are more successful in supporting breeding pairs of this species (Musil 2006). The cyclical removal of adult fish from an artificial waterbody (gravel pit) in the UK resulted in an increase in invertebrate food availability and increases in the growth of submerged aquatic macrophytes, which in turn led to a increase in brood use of the habitat by the species and increased species survival (Giles 1994). The removed fish (dead or alive) were sold to generate funds (Giles 1994).

Threats
The species is threatened by habitat degradation due to oil pollution, drainage, peat-extraction and changing land management practices (Grishanov 2006) (e.g. decreased grazing and mowing in meadows leading to scrub over-growth (Grishanov 2006) and agricultural intensification (Kear 2005a)) in breeding areas (Kear 2005a, Grishanov 2006). It also suffers decreased reproductive success as a result of disturbance from increased recreational use of inland waterbodies (Kear 2005a), machinery noise from urban development (Marsden 2000), hunting (Evans and Day 2002) and predation by American mink Neovison vison on islands (Nordstrom et al. 2002). The species is susceptible to avian influenza so may be threatened by future outbreaks of the virus (Melville and Shortridge 2006). Utilisation The species is hunted for sport Denmark (Bregnballe et al. 2006), Italy (Sorrenti et al. 2006) and Iran (Balmaki and Barati 2006), and is hunted commercially in Iran (Balmaki and Barati 2006). The eggs of this species were (and possibly still are) harvested in Iceland (Gudmundsson 1979).

References
Balmaki, B.; Barati, A. 2006. Harvesting status of migratory waterfowl in northern Iran: a case study from Gilan Province. In: Boere, G.; Galbraith, C., Stroud, D. (ed.), Waterbirds around the world, pp. 868-869. The Stationary Office, Edinburgh, UK.

Brazil, M. 2009. Birds of East Asia: eastern China, Taiwan, Korea, Japan, eastern Russia. Christopher Helm, London.

Bregnballe, T.; Noer, H.; Christensen, T. K.; Clausen, P.; Asferg, T.; Fox, A. D.; Delany, S. 2006. Sustainable hunting of migratory waterbirds: the Danish approach. In: Boere, G.; Galbraith, C., Stroud, D. (ed.), Waterbirds around the world, pp. 854-860. The Stationary Office, Edinburgh, UK.

del Hoyo, J.; Elliot, A.; Sargatal, J. 1992. Handbook of the Birds of the World, vol. 1: Ostrich to Ducks. Lynx Edicions, Barcelona, Spain.

Delany, S.; Scott, D. 2006. Waterbird population estimates. Wetlands International, Wageningen, The Netherlands.

Evans, D. M.; Day, K. R. 2002. Hunting disturbance on a large shallow lake: the effectiveness of waterfowl refuges. Ibis 144(1): 2-8.

Flint, V. E.; Boehme, R. L.; Kostin, Y. V.; Kuznetsov, A. A. 1984. A field guide to birds of the USSR. Princeton University Press, Princeton, New Jersey.

Giles, N. 1994. Tufted Duck (Aythya fuligula) habitat use and brood survival increases after fish removal from gravel pit lakes. Hydrobiologia 279/280: 387-392.

Grishanov, D. 2006. Conservation problems of migratory waterfowl and shorebirds and their habitats in the Kaliningrad region of Russia. In: Boere, G.; Galbraith, C., Stroud, D. (ed.), Waterbirds around the world, pp. 356. The Stationary Office, Edinburgh, UK.

Gudmundsson, F. 1979. The past status and exploitation of the Myvatn waterfowl populations. Oikos 32((1-2)): 232-249.

Kear, J. 2005. Ducks, geese and swans volume 2: species accounts (Cairina to Mergus). Oxford University Press, Oxford, U.K.

Madge, S.; Burn, H. 1988. Wildfowl. Christopher Helm, London.

Marsden, S. J. 2000. Impact of Disturbance on Waterfowl Wintering in a UK Dockland Redevelopment Area. Environmental Management 26(2): 207-213.

Marsden, S. J.; Bellamy, G. S. 2000. Microhabitat characteristics of feeding sites used by diving duck Aythya wintering on the grossly polluted Manchester Ship Canal, UK. Environmental Conservation 27(3): 278-283.

Melville, D. S.; Shortridge, K. F. 2006. Migratory waterbirds and avian influenza in the East Asian-Australasian Flyway with particular reference to the 2003-2004 H5N1 outbreak. In: Boere, G.; Galbraith, C., Stroud, D. (ed.), Waterbirds around the world, pp. 432-438. The Stationary Office, Edinburgh, UK.

Musil, P. 2006. A review of the effects of intensive fish production on waterbird breeding populations. In: Boere, G.; Galbraith, C., Stroud, D. (ed.), Waterbirds around the world, pp. 520-521. The Stationary Office, Edinburgh, UK.

Nordström, M.; Högmander, J.; Nummelin, J.; Laine, J.; Laanetu, N.; Korpimäki, E. 2002. Variable responses of waterfowl breeding populations to long-term removal of introduced American mink. Ecography 25: 385-394.

Scott, D. A.; Rose, P. M. 1996. Atlas of Anatidae populations in Africa and western Eurasia. Wetlands International, Wageningen, Netherlands.

Snow, D. W.; Perrins, C. M. 1998. The Birds of the Western Palearctic vol. 1: Non-Passerines. Oxford University Press, Oxford.

Sokolov, L. V.; Gordienko, N. S. 2008. Has recent climate warming affected the dates of bird arrival to the Il'men Reserve in the Southern Urals? Russian Journal of Ecology 39: 56-62.

Sorrenti, M.; Carnacina, L.; Radice, D.; Costato, A. 2006. Duck harvest in the Po delta, Italy. In: Boere, G.; Galbraith, C., Stroud, D. (ed.), Waterbirds around the world, pp. 864-865. The Stationary Office, Edinburgh, UK.

Winfield, D. K.; Winfield, I. J. 1994. Possible competitive interactions between overwintering tufted duck (Aythya fuligula (L.)) and fish populations of Lough Neagh, Northern Ireland: evidence from diet studies. Hydrobiologia 279/280: 377-386.

Further web sources of information
Detailed species account from Birds in Europe: population estimates trends and conservation status (BirdLife International 2004)

Detailed species account from Birds in Europe: population estimates, trends and conservation status (BirdLife International 2004)

Explore HBW Alive for further information on this species

Search for photos and videos, and hear sounds of this species from the Internet Bird Collection

Text account compilers
Ekstrom, J., Butchart, S., Malpas, L.

IUCN Red List evaluators
Butchart, S., Symes, A.

Recommended citation
BirdLife International (2014) Species factsheet: Aythya fuligula. Downloaded from http://www.birdlife.org on 22/12/2014. Recommended citation for factsheets for more than one species: BirdLife International (2014) IUCN Red List for birds. Downloaded from http://www.birdlife.org on 22/12/2014.

This information is based upon, and updates, the information published in BirdLife International (2000) Threatened birds of the world. Barcelona and Cambridge, UK: Lynx Edicions and BirdLife International, BirdLife International (2004) Threatened birds of the world 2004 CD-ROM and BirdLife International (2008) Threatened birds of the world 2008 CD-ROM. These sources provide the information for species accounts for the birds on the IUCN Red List.

To provide new information to update this factsheet or to correct any errors, please email BirdLife

To contribute to discussions on the evaluation of the IUCN Red List status of Globally Threatened Birds, please visit BirdLife's Globally Threatened Bird Forums.

Additional resources for this species

ARKive species - Tufted duck (Aythya fuligula) 0

Key facts
Current IUCN Red List category Least Concern
Family Anatidae (Ducks, Geese, Swans)
Species name author (Linnaeus, 1758)
Population size mature individuals
Population trend Stable
Distribution size (breeding/resident) 14,400,000 km2
Country endemic? No
Links to further information
- Additional Information on this species