| Location | Uganda, Kabale,Kisoro,Rukungiri |
| Central coordinates | 29o 40.00' East 1o 3.00' South |
| IBA criteria | A1, A2, A3 |
| Area | 32,700 ha |
| Altitude | 1,190 - 2,607m |
| Year of IBA assessment | 2001 |
Ornithological information See Box and Tables 2 and 3 for key species. The National Park has been extensively surveyed for birds. Its checklist currently totals 347 species. Mubwindi swamp is home to Bradypterus graueri. Indicator pumilio is only known in Uganda, with certainty, from this locality. Some of the species endemic to the Albertine Rift, such as Pseudocalyptomena graueri, Muscicapa lendu and Cryptospiza shelleyi, have limited distributions elsewhere in their range. The park also holds Zoothera oberlaenderi, one of the six species of the Eastern DR Congo lowlands Endemic Bird Area (EBA 107). The northern sector is especially rich in species of the Guinea–Congo Forests biome.
Site description The park lies on the edge of the Albertine (or Western) Rift Valley and occupies the highest block of the Kigezi highlands, also known as the Rukiga highlands. It is located on Uganda’s border with the Democratic Republic of Congo (DRC). The topography of the park is extremely rugged and much dissected, especially in the higher southern sector. The only flat area of any size is Mubwindi swamp (c.80 ha). The remainder (almost all forested) consists of narrow, very steep-sided valleys bounded by hill crests, spanning altitudes between 1,400 m in the northern sector and 2,600 m in the south.The forest is one of the largest in East Africa that contains both medium-altitude and montane forest in a continuum. As a result of its wide altitudinal range and relatively large size, and its probable role as a Pleistocene refugium, the species-richness of the forest is extremely high, for both flora and fauna. This forest is believed to be a mere remnant of a very large forest which once covered much of western Uganda, Rwanda, Burundi and eastern DRC. Today, Bwindi is an ecological island surrounded by one of the highest human population densities in Africa (100–450 individuals/km²). Immediately beyond its borders, there is virtually no natural forest remaining and much of the land is intensively cultivated.The Institute of Tropical Forest Conservation, which belongs to Mbarara University of Science and Technology, is based at Ruhija. The park is host to considerable ecotourist activities, particularly gorilla-tracking and birdwatching.
| Species | Season | Period | Population estimate | Quality of estimate | IBA Criteria | IUCN Category |
|---|---|---|---|---|---|---|
| Handsome Francolin Francolinus nobilis | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Mountain Buzzard Buteo oreophilus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Cassin's Hawk-eagle Aquila africanus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-spotted Flufftail Sarothrura pulchra | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Afep Pigeon Columba unicincta | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Dusky Turtle-dove Streptopelia lugens | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black-collared Lovebird Agapornis swindernianus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grey Parrot Psittacus erithacus | resident | 1998 | present [units unknown] | - | A3 | Not Recognised |
| Ruwenzori Turaco Ruwenzorornis johnstoni | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Dusky Long-tailed Cuckoo Cercococcyx mechowi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Olive Long-tailed Cuckoo Cercococcyx olivinus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Fraser's Eagle-owl Bubo poensis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Red-chested Owlet Glaucidium tephronotum | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black-shouldered Nightjar Caprimulgus nigriscapularis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Montane Nightjar Caprimulgus poliocephalus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Ruwenzori Nightjar Caprimulgus ruwenzorii | resident | 1998 | present [units unknown] | - | A2 | Least Concern |
| Scarce Swift Schoutedenapus myoptilus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Bar-tailed Trogon Apaloderma vittatum | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Blue-throated Roller Eurystomus gularis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black Bee-eater Merops gularis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Cinnamon-chested Bee-eater Merops oreobates | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Forest Woodhoopoe Phoeniculus castaneiceps | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black-and-white-casqued Hornbill Bycanistes subcylindricus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Brown-cheeked Hornbill Bycanistes cylindricus | resident | 1998 | present [units unknown] | - | A3 | Vulnerable |
| Speckled Tinkerbird Pogoniulus scolopaceus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Western Green-tinkerbird Pogoniulus coryphaeus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-throated Tinkerbird Pogoniulus subsulphureus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-spotted Barbet Buccanodon duchaillui | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Hairy-breasted Barbet Tricholaema hirsuta | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-billed Barbet Trachyphonus purpuratus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Cassin's Honeyguide Prodotiscus insignis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Willcocks's Honeyguide Indicator willcocksi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Dwarf Honeyguide Indicator pumilio | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Near Threatened |
| Tullberg's Woodpecker Campethera tullbergi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Buff-spotted Woodpecker Campethera nivosa | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Brown-eared Woodpecker Campethera caroli | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Golden-crowned Woodpecker Thripias xantholophus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| African Green Broadbill Pseudocalyptomena graueri | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Vulnerable |
| Green-breasted Pitta Pitta reichenowi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| African Shrike-flycatcher Megabyas flammulatus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Ruwenzori Batis Batis diops | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Chestnut Wattle-eye Platysteira castanea | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Jameson's Wattle-eye Platysteira jamesoni | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Chestnut-bellied Helmet-shrike Prionops caniceps | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grey-green Bush-shrike Telophorus bocagei | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Doherty's Bush-shrike Telophorus dohertyi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Lagden's Bush-shrike Malaconotus lagdeni | resident | - | present [units unknown] | - | A1 | Near Threatened |
| Sooty Boubou Laniarius leucorhynchus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Mountain Boubou Laniarius poensis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grey Cuckooshrike Coracina caesia | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Petit's Cuckooshrike Campephaga petiti | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Western Black-headed Oriole Oriolus brachyrhynchus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black-tailed Oriole Oriolus percivali | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black-headed Paradise-flycatcher Terpsiphone rufiventer | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Dusky Crested-flycatcher Elminia nigromitrata | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-bellied Crested-flycatcher Elminia albiventris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-tailed Crested-flycatcher Elminia albonotata | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Dusky Tit Parus funereus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Stripe-breasted Tit Parus fasciiventer | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Chubb's Cisticola Cisticola chubbi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Carruthers's Cisticola Cisticola carruthersi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Collared Apalis Oreolais ruwenzorii | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Black-faced Apalis Apalis personata | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Buff-throated Apalis Apalis rufogularis | resident | 1998 | present [units unknown] | - | A3 | Not Recognised |
| Chestnut-throated Apalis Apalis porphyrolaema | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-browed Camaroptera Camaroptera superciliaris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Olive-green Camaroptera Camaroptera chloronota | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Shelley's Greenbul Andropadus masukuensis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grey Greenbul Andropadus gracilis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Plain Greenbul Andropadus curvirostris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Eastern Mountain Greenbul Andropadus nigriceps | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Honeyguide Greenbul Baeopogon indicator | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Joyful Greenbul Chlorocichla laetissima | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Toro Olive Greenbul Phyllastrephus hypochloris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Icterine Greenbul Phyllastrephus icterinus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Common Bristlebill Bleda syndactylus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Red-tailed Bulbul Criniger calurus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-spotted Nicator Nicator chloris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Neumann's Warbler Hemitesia neumanni | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| White-winged Scrub-warbler Bradypterus carpalis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grauer's Swamp-warbler Bradypterus graueri | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Endangered |
| Mountain Flycatcher-warbler Chloropeta similis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Green Hylia Hylia prasina | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Red-faced Woodland-warbler Phylloscopus laetus | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Grauer's Warbler Graueria vittata | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Turner's Eremomela Eremomela turneri | resident | 1998 | present [units unknown] | - | A3 | Endangered |
| Green Crombec Sylvietta virens | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-browed Crombec Sylvietta leucophrys | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Scaly-breasted Illadopsis Illadopsis albipectus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Brown Illadopsis Illadopsis fulvescens | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Mountain Illadopsis Illadopsis pyrrhoptera | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grey-chested Babbler Kakamega poliothorax | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| African Hill Babbler Pseudoalcippe abyssinica | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Capuchin Babbler Phyllanthus atripennis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Purple-headed Glossy-starling Lamprotornis purpureiceps | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Sharpe's Starling Cinnyricinclus sharpii | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Waller's Starling Onychognathus walleri | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Slender-billed Starling Onychognathus tenuirostris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Stuhlmann's Starling Poeoptera stuhlmanni | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Narrow-tailed Starling Poeoptera lugubris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-tailed Ant-thrush Neocossyphus poensis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Kivu Ground-thrush Zoothera tanganjicae | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Not Recognised |
| Forest Ground-thrush Zoothera oberlaenderi | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Near Threatened |
| Black-eared Ground-thrush Zoothera camaronensis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Red-throated Alethe Alethe poliophrys | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| White-tailed Alethe Alethe diademata | resident | 1998 | present [units unknown] | - | A3 | Not Recognised |
| White-starred Robin Pogonocichla stellata | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Equatorial Akalat Sheppardia aequatorialis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-bellied Robin-chat Cossyphicula roberti | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Archer's Robin-chat Cossypha archeri | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Blue-shouldered Robin-chat Cossypha cyanocampter | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-eyed Black Flycatcher Melaenornis ardesiacus | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| White-eyed Slaty Flycatcher Dioptrornis fischeri | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Sooty Flycatcher Muscicapa infuscata | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Chapin's Flycatcher Muscicapa lendu | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Vulnerable |
| Dusky-blue Flycatcher Muscicapa comitata | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Cassin's Grey Flycatcher Muscicapa cassini | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Grey-throated Tit-flycatcher Myioparus griseigularis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Scarlet-tufted Sunbird Anthreptes fraseri | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Green Sunbird Anthreptes rectirostris | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Little Green Sunbird Nectarinia seimundi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Blue-headed Sunbird Nectarinia alinae | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Blue-throated Brown Sunbird Nectarinia cyanolaema | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Green-throated Sunbird Nectarinia rubescens | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Northern Double-collared Sunbird Nectarinia preussi | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Regal Sunbird Nectarinia regia | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Purple-breasted Sunbird Nectarinia purpureiventris | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Bronze Sunbird Nectarinia kilimensis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Superb Sunbird Nectarinia superba | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Baglafecht Weaver Ploceus baglafecht | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Black-billed Weaver Ploceus melanogaster | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Strange Weaver Ploceus alienus | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Vieillot's Black Weaver Ploceus nigerrimus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-mantled Weaver Ploceus tricolor | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Brown-capped Weaver Ploceus insignis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Red-headed Malimbe Malimbus rubricollis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Woodhouse's Antpecker Parmoptila woodhousei | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-breasted Negrofinch Nigrita fusconotus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| White-collared Oliveback Nesocharis ansorgei | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Red-faced Crimson-wing Cryptospiza reichenovii | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Abyssinian Crimson-wing Cryptospiza salvadorii | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Dusky Crimson-wing Cryptospiza jacksoni | resident | 1998 | present [units unknown] | - | A2, A3 | Least Concern |
| Shelley's Crimson-wing Cryptospiza shelleyi | resident | 1998 | present [units unknown] | - | A1, A2, A3 | Vulnerable |
| Dusky Twinspot Euschistospiza cinereovinacea | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Swee Waxbill Estrilda melanotis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Yellow-browed Citril Serinus frontalis | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Streaky Seedeater Serinus striolatus | resident | 1998 | present [units unknown] | - | A3 | Not Recognised |
| Thick-billed Seedeater Serinus burtoni | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| Oriole Finch Linurgus olivaceus | resident | 1998 | present [units unknown] | - | A3 | Least Concern |
| IUCN habitat | Habitat detail | Extent (% of site) |
|---|---|---|
| Artificial landscapes (terrestrial) | 1% | |
| Forest | 97% |
| Land-use | Extent (% of site) |
|---|---|
| nature conservation and research | - |
| tourism/recreation | - |
| water management | - |
| other | - |
| Notes: Collection of forest products. | |
Other biodiversity Eight species of tree are known only from this forest in Uganda. Among threatened mammals, the forest contains about 300 individuals of Gorilla gorilla beringei (CR), roughly half of the world population of this subspecies, as well as Loxodonta africana (EN), Pan troglodytes (EN) and Cercopithecus l’hoesti (LR/nt).
Management considerations This forest was originally gazetted in 1932 as Kasatoro Crown Forest (south sector, 181 km²) and Kayonza Crown Forest (north sector, 26 km²). In 1942, the two forests were combined and additional forest was added to establish the Impenetrable Crown Forest (324 km²). About 26 km² were excised from the reserve in 1948, and it was re-gazetted the Impenetrable Central Forest Reserve (298 km²). These gazettements were consolidated in the Laws of Uganda in 1951. Additional small excisions were made in 1958 in order to shorten the boundaries and make them easier to maintain and patrol. In 1961, two local Forest Reserves bordered the Impenetrable; the Bikingi Local Forest Reserve (7.6 km²) in the south sector and the Ishasha Local Forest Reserve (13.8 km²) in the north sector. In 1966, these were incorporated into the Impenetrable Central Forest Reserve (320.8 km²). The Impenetrable was gazetted as an Animal Sanctuary in 1961 with boundaries the same as those of the Impenetrable Central Forest Reserve, which was then gazetted as a National Park in 1991. This change in national status reflects the growing recognition of Bwindi Impenetrable Forest as a vital refuge for some of Uganda’s rarest and most unique flora and fauna.Bwindi Impenetrable Forest is the largest remaining fragment of natural forest in south-western Uganda, with exceptional species-richness. It is an important water catchment, being the source of rivers flowing to Lakes Edward, Mutanda and Bunyonyi, and to the drier agricultural land to the north and west. For these reasons, the Bwindi Impenetrable Forest has been declared a World Heritage Site, together with Rwenzori Mountains National Park. Owing to the size of the forest, it may have an important stabilizing effect on the microclimate of the area.Conservation efforts in Bwindi have historically faced a variety of obstacles, mainly derived from the conflicts of interest over land usage, specifically the desire, on the part of the community members, to utilize park resources as they had done traditionally. Rapid population growth and high population densities in the region, combined with poor agricultural practices, poorly developed local and national transport systems and/or techniques for natural-resource conservation, perpetuate local community dependence upon forest resources, although this is changing. Policies and practices of the National Park also fostered conservation conflicts. Local communities were historically excluded from the decision-making, planning and management of the forest, which contributed to resentment and hostility by local people towards park authorities. All this culminated in illegal activities and disinterest by local communities in the park. In addition, local people incur high costs in the form of loss of access to resources and crop depredation by wildlife but, in the past, received little tangible benefit from the park establishment. However, the majority of illegal activities have been brought under control and considerable efforts are being made by organizations such as CARE (Uganda) and the Mgahinga and Bwindi Impenetrable Forest Conservation Trust to assist local communities to adopt sustainable resource-use practices. By 2000, these measures were beginning to have a significant effect. Gorilla-based tourism is most lucrative for the Uganda Wildlife Authority, and part of the proceeds are now ploughed back into surrounding communities.
References Butynski (1984), Gubelman, (1995), Harcourt et al. (1983), Howard (1991), Kalina and Butynski (1996), Keith et al. (1969), Langdale-Brown et al. (1964), Leggat and Osmaston (1961), Marks and Gnoske (1997), Mwambu (1999).
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Recommended citation BirdLife International (2013) Important Bird Areas factsheet: Bwindi Impenetrable National Park. Downloaded from http://www.birdlife.org on 19/06/2013
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